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Preheat the oven to 190°C (170°C fan) / Gas Mark 5 and set aside a 23x33cm baking dish.
Bring a large saucepan of salted water to the boil.
Add the dried macaroni and cook until just tender (al dente), following the packet instructions, then drain and set aside.
In a large saucepan over a medium heat, melt the unsalted butter.
Stir in the plain flour and whisk constantly for 1–2 minutes to form a roux.
Gradually add the whole milk, whisking continuously, until the sauce is smooth and lump-free.
Continue to cook, stirring frequently, for about 5–7 minutes, until the sauce coats the back of a spoon.
Remove from the heat and stir in the grated mature cheddar cheese and grated mozzarella until fully melted.
Season the sauce with the salt, freshly ground black pepper and a pinch of ground nutmeg.
Add the cooked pasta to the cheese sauce and stir gently until all the tubes are evenly coated.
Spoon the mixture into the prepared dish. If using, sprinkle the Panko breadcrumbs evenly over the top.
Bake for 25–30 minutes, until bubbling around the edges and the top is golden brown and crisp.
Leave to rest for 5–10 minutes before serving.